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MUSEUM D'HISTOIRE NATURELLE de la Ville de Genève

tome 110 fascicule 3 2003

g kl GENEVE SEPTEMBRE 2003 ISSN 0035 - 418 X

REVUE SUISSE DE ZOOLOGIE

SWISS JOURNAL OF ZOOLOGY

REVUE SUISSE DE ZOOLOGIE

TOME 110—FASCICULE 3

Publication subventionnée par: ACADÉMIE SUISSE DES SCIENCES NATURELLES ASSN VILLE DE GENÈVE SOCIÉTÉ SUISSE DE ZOOLOGIE

VOLKER MAHNERT Directeur du Muséum d'histoire naturelle de Genève

CHARLES LIENHARD Chargé de recherche au Muséum d'histoire naturelle de Genève

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Les manuscrits sont soumis à des experts d'institutions suisses ou étrangères selon le sujet étudié.

La préférence sera donnée aux travaux concernant les domaines suivants: biogéo- graphie, systématique, évolution, écologie, éthologie, morphologie et anatomie comparée, physiologie.

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MUSEUM D'HISTOIRE NATURELLE 1211 GENÈVE 6

Internet: http://www.ville-ge.ch/musinfo/mhng/page/rsz.htm

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ANNALES

de la

SOCIÉTÉ SUISSE DE ZOOLOGIE

et du

MUSEUM D'HISTOIRE NATURELLE de la Ville de Genéve

tome 110 fascicule 3 2003

g f EI GENEVE SEPTEMBRE 2003 ISSN 0035 - 418 X

REVUE SUISSE DE ZOOLOGIE

SWISS JOURNAL OF ZOOLOGY

REVUE SUISSE DE ZOOLOGIE

TOME 110—FASCICULE 3

Publication subventionnée par: ACADÉMIE SUISSE DES SCIENCES NATURELLES ASSN VILLE DE GENÈVE SOCIÉTÉ SUISSE DE ZOOLOGIE

VOLKER MAHNERT Directeur du Muséum d'histoire naturelle de Genève

CHARLES LIENHARD Chargé de recherche au Muséum d'histoire naturelle de Genève

Comité de lecture

Il est constitué en outre du président de la Société suisse de Zoologie, du directeur du Muséum de Genève et de représentants des instituts de zoologie des universités suisses.

Les manuscrits sont soumis à des experts d'institutions suisses ou étrangères selon le sujet étudié.

La préférence sera donnée aux travaux concernant les domaines suivants: biogéo- graphie, systématique, évolution, écologie, éthologie, morphologie et anatomie comparée, physiologie.

Administration

MUSEUM D'HISTOIRE NATURELLE 1211 GENÈVE 6

Internet: http://www.ville-ge.ch/musinfo/mhng/page/rsz.htm

PRIX DE L’ABONNEMENT:

SUISSE Fr. 225.— UNION POSTALE Fr. 230.—

(en francs suisses)

Les demandes d’abonnement doivent être adressées à la rédaction de la Revue suisse de Zoologie, Muséum d'histoire naturelle, C.P. 6434, CH-1211 Genève 6, Suisse

REVUE SUISSE DE ZOOLOGIE 110 (3): 453-481; septembre 2003

Oribatids from Switzerland VII (Acari: Oribatida: Ptyctima). (Acarologica Genavensia CII)

Sandor MAHUNKA & Luise MAHUNKA-PAPP Department of Zoology, Hungarian Natural History Museum, Baross utca 13, H-1088 Budapest, Hungary.

Oribatids from Switzerland VIII (Acari: Oribatida: Ptyctima). (Acarologica Genavensia CII). - Earlier published and newly determined Swiss moss mites belonging to the “ptychoid” groups are listed, discussed or described. From the 20 named species in the literature 10 are confirmed after critical discussion. New identifications of a total of 42 taxa are given, 3 of which (Phthiracarus besuchetianus sp. n., Steganacarus (S.) anten- natus Sp. n. and Steganacarus (S.) schweizeri sp. n.) are new to science and 30 new for the Fauna of Switzerland. Some new or additional morpho- logical data as well as some notes on the taxonomy and distribution of this group are given.

Key-words: Acari - Oribatida - Ptyctima - taxonomy - new species - Switzerland.

INTRODUCTION

The present paper, the 8th part of the series (concerning our project, started in July 1983 see Mahunka & Mahunka-Papp, 2001), proposes to discuss our results gained on the ptychoid groups of the Swiss Fauna comprising the already published references and our recent results.

New identifications of 42 species and subspecies belonging to the families Phthiracaridae, Steganacaridae, Euphthiracaridae, Oribotritiidae and Mesoplophori- dae, found on the territory of Switzerland, are given. Among them 3 are new for science and 30 are new for the Fauna of Switzerland. Some rare species are recorded and the locality data for all our identifications are given. Morphological details, corrections and the interpretation of some species are also given.

Unfortunately, the early data are mostly uncertain because there is perhaps no other oribatid group in recent decades which has suffered so many changes than this one due to the arduous work of Parry/Kamill (Parry, 1979; Kamill & Baker, 1980; Kamill, 1981), Niedbata (Niedbata, 1986a; Niedbata, 1986b; Niedbata, 1992) Bernini (e.g. Bernini & Avanzati, 1986) and others (see Balogh & Mahunka, 1983; Balogh & Balogh, 1992). The 20 named species (under the same name different authors under- stand different species) known from the earlier literature are critically discussed, the existing slides partly revised and 10 of them accepted as members of the Swiss Fauna.

Manuscript accepted 10.12.2002

454 S. MAHUNKA & L. MAHUNKA-PAPP

This contribution follows the system of Niedbata (1986a, 1992) with some slight modifications, also the terminology of the morphological descriptions is modified as in our previous papers (e.g. Mahunka, 1990). Contrary to our earlier prac- tice we introduce a significant change in systematics, 1.e. the abandonment of the sub- genus Archiphthiracarus Balogh & Mahunka, 1979, heretofore retained for practical purposes. Since there is not a single species of the “Archiphthiracarus” group (5 pairs of anoadanal setae) known in which a complete reduction of two pairs of the adanal setae could be observed (there are always 1-2 partially vestigial pairs of setae), we see no reason to conserve this subgenus. Presently the species P. globosus (C. L. Koch, 1841) displays both variations, consequently, the concept of Niedbata (1992) should be followed.

HITHERTO PUBLISHED SPECIES OF SWISS PTYCTIMA

HISTORY

This group has been studied and results published by Schweizer (1922, 1948, 1956), Borcard (1988, 1991) and Mahunka (1993a, 1993b).

The following chronological list recapitulates all recorded species with the published name and the actual valid name or, in the case of misidentifications, the correct species.

SCHWEIZER, 1922

Hoploderma magnum (Nic.) = Steganacarus magnus (Nicolet, 1855) Hoploderma dasypus (Ant. Dugès) = species inquirenda

Hoploderma globosum (C. L. Koch) = Phthiracarus globosus (C. L. Koch, 1841) Hoploderma striculum (C. L. Koch) = Atropacarus striculus (C. L. Koch, 1835) Phthiracarus arduus (C. L. Koch) = Rhysotritia ardua (C. L. Koch, 1841)

SCHWEIZER, 1948

Hoploderma laevigatum (C. L. Koch) = ?

Hoploderma spinosum (Sellnick) = ?

Hoploderma magnum (Nic.) = Steganacarus magnus (Nicolet, 1855) Phthiracarus pavidus (Berl.) = ?

Phthiracarus piger (Scopoli) = species inquirenda

Phthiracarus globosus (C. L. Koch)

SCHWEIZER, 1956

Phthiracarus borealis (Trhgd.) = ?

Phthiracarus crenophilus Willmann, 1951

Hoploderma laevigatum (C. L. Koch) = ?

Hoploderma spinosum (Sellnick) = Steganacrus antennatus sp. n.

Hoploderma striculum (C. L. Koch) = Atropacarus striculus (C. L. Koch, 1835) Hoploderma pavidum Berlese, 1913 =?

Hoploderma clavigerum Berlese, 1903 = ?

Oribotrita [sic!] nuda (Berlese) = Mesotritia nuda (Berlese, 1887)

Oribotrita [sic!] canestrinii Berlese [sic!] = Rhysotritia ardua (C. L. Koch, 1841)

BORCARD, 1991

Hoplophthiracarus pavidus (Berlese) 1913 = ? Hoplophthiracarus vanderhammeni Niedbata, 1991

Phthiracarus cf. tardus Forsslund 1956 = Phthiracarus cf. longulus (C. L. Koch, 1841)

ORIBATIDS FROM SWITZERLAND VIII 455

Phthiracarus sp. À = ?

Phthiracarus sp. B =?

Phthiracarus sp. C =?

Steganacarus herculaneus Willmann, 1951

Steganacarus striculus (C. L. Koch) 1836 = Atropacarus striculus (C. L. Koch, 1835) Rhysotritia ardua (C. L. Koch) 1841

MAHUNKA, 1993a Helvetacarus genavensis Mahunka, 1993

MAHUNKA, 1993b

Rhysotritia a. ardua C. L. Koch, 1841

Atropacarus wandae Niedbata, 1981

Phthiracarus (Archiphthiracarus) bryobius Jacot, 1930 = Phthiracarus bryobius Jacot, 1930

Phthiracarus (Phthiracarus) globosus (C. L. Koch, 1841) = Phthiracarus globosus (C. L. Koch, 1841)

CRITICAL CONSIDERATIONS

GENERAL REMARKS ON SCHWEIZER ’S PUBLICATIONS

It was Schweizer who published the most numerous data on ptychoid groups similarly to other superfamilies. A significant part of the species mentioned by him could be studied due to the help of the late custodian of the Naturhistorisches Museum Basel, Dr C. Bader, our good friend (see also Bader, 1969). Unfortunately, the material is in poor condition, which is partly due to the permanent slides (which, for obvious reasons, could not be remounted), and partly to the fact that some of the slides suffered heavy damage, so could not be studied at all, and also because Schweizer out of necessity amassed sometimes 10-20 specimens on a single slide. These slides occasionally include specimens belonging to 5-6 genera, which he obviously knew. Furthermore, he also committed mistakes in the grouping of specimens, at the species or even at the generic level, in such cases the specimens of 2-3 different Phthiracarus species are found side by side and, to further aggravate the problem, under one species name.

So partly for these and some other reasons we could not study all his available slides, so his identifications in the collection need further, thorough investigation.

Schweizer listed a total of 14 “ptychoid” species in his works. On the basis of his drawings and slides this number is reduced to six as belonging to the fauna of Switzerland. Quite obviously he understood different species under other names. We accept the following six species of Schweizer to be members of the Swiss Fauna:

Phthiracarus crenophilus Willmann, 1951

Phthiracarus globosus (C. L. Koch, 1841)

Steganacarus magnus (Nicolet, 1855)

Atropacarus striculus (C. L. Koch, 1835)

Rhysotritia ardua (C. L. Koch, 1841) Mesotritia nuda (Berlese, 1887)

SCHWEIZER, 1922

In his first work he mentioned five species, which had been identified most likely by using the work of Michael (1898). However, no voucher specimens have been

456 S. MAHUNKA & L. MAHUNKA-PAPP

found in the collection of the Naturhistorisches Museum, Basel. In all probability, from among the five species Hoploderma magnum, Hoploderma globosum, Hoploderma striculum and Phthiracarus arduus were correctly determined, although H. magnum was interpreted in his subsequent works as a Phthiracarus species. In the case of H. dasypus it is best to accept the standpoint of Niedbata (1992), i.e. the status of the species is wholly uncertain, so the identification of Schweizer cannot be evaluated at all.

Generally Schweizer has not made reference to his earlier data, so he makes no reference to his identifications of 1922 in his subsequent papers of 1948 or 1956. Indeed he makes no particular reference to his species listed in 1948 in his paper published in 1956, although it appears quite obvious that he has these 1948 species in mind. Part of the recovered slides also supports this suggestion.

SCHWEIZER, 1948

This work listed six species. However, only two species (H. magnum, H. glo- bosum) were mentioned in 1922, but three (H. laevigatum, H. spinosum and H. pavidum) are repeated in 1956. Oddly he does not mention the species H. magnum, Phthiracarus piger and P. globosus in 1956.

Of the three species H. magnum and P. globosus may be accepted as good iden- tifications, however, P. piger Scopoli, 1763 is a species inquirenda as is the already mentioned P. dasypus (see Niedbata, 1992: 3). In connection with this species we cannot accept the view of Berg et al. (1990) that it is P. piger, because the neotypes were designated by van der Hammen (1963, 1964) upon the redescription of P. laevi- gatus and P. nitens (= P. piger sensu Berg et al., 1990) (which was not made in the work of Berg er al., 1990). Consequently, the rule of the ICZN should be applied, according to which the name of the first specialist ($ 24) revising the name should be used.

We have not found a single specimen of the species H. globosus. The material of P. piger which we had received contained only slides: N 786 (“Mtr. Pr. 1, 13. VII. 30, V. d. Botsch, 2100 m. Wurzelgefl.”)!, N 1426 and N 1427 (“No. 8, Scarl, Lawiner, 12. VII. 1932, v. Holz’). The species name of piger on all three slides was crossed out, most likely by Schweizer himself: One slide bears the name borealis, the other two crenophilus, displaying identical hand-writing as for the generic name. We have established that the slide bearing borealis includes specimens of the species P. bryobius Jacot, 1930, while the other two bearing the name P. crenophilus Willmann, 1951 had been correctly identified.

SCHWEIZER, 1956

The species of Hoploderma pavidum (Berlese, 1913) was misunderstood by Schweizer. The slides (N 668, N 1288 and N 1745) said to contain these specimens, in fact, display a so far unidentified Steganacarus species. Consequently, H. pavidum as referred to by Schweizer should be deleted from the fauna of Switzerland.

! Orthography is reported just as on the labels of the slides.

ORIBATIDS FROM SWITZERLAND VIII 457

We have also found the slide N 583 (“Mtr. Pr. 6, 20. VII. 29, Val Cluozza, Valetta, Bachmoos 1900 m.”) containing the species Hoploderma laevigatum as listed and figured by Schweizer. In fact, it has nothing to do with P. laevigatum itself. Quite unequivocally it may be stated again that it is an unidentified Steganacarus species. On this very same slide were also embedded the species P. spadix Niedbata, 1983 and another Phthiracarus species. So P. laevigatus should also be deleted from the fauna list of Switzerland, and this applies for the previous publications too.

The species Hoploderma striculum (C.L. Koch, 1835) was recorded by Schweizer several times (1922, 1956). We have found it on a number of slides, as for example N 1267, 1268 and 1274 (“Mtr. Pr. XII, Punt Perif, 23. VII. 1930, Moos, Wasser-Kante d. Spöl.”), and the well-preserved specimens confirm a correct iden- tification.

We have found several specimens of the species Hoploderma clavigerum as identified by Schweizer on slides N 1819 and N 1820 (“Mtr. Pr. XV, Scarl, 2000 m, 24. VII. 1932, in Mull von Bergföhren-Nadeln, Sammelpräparat”), so it is certain that the species had been identified by Schweizer on the basis of these specimens. However, they belong to a so far undescribed Steganacarus species, since the chaetotaxy of leg IV is clearly discernible. This species again should be deleted from the list of the Swiss fauna.

The Schweizer identified Hoploderma spinosum has also been recovered in the collection. This species is however a new one which is described in this contribution in the Steganacarus genus (S. antennatus sp. n.). S. spinosus (Sellnick, 1920) should also be deleted from the Swiss fauna list.

In his work of 1956 Schweizer also identified two Oribotrita [sic!] species. Oribotritia (= Mesotritia) nuda (Berlese, 1887) was represented only on one slide (N 891) with the following data: “N 53, 17. VII. 1930 Stavel-chod, 2100, u. Holz”. Although the examined specimen is well preserved, without remounting a true identi- fication is impossible, for many significant features cannot be studied. Nevertheless, we consider the identification to be correct.

The specimens referring to the species of Oribitritia canestrinii (Michael, 1898) were located and studied on the slides marked by H 18 226 and H 18 231. The specimens deriving from the collecting locality of “Scanfs 1700 m. Heide mit Wach- older + Erikas” were unequivocally identified with Rhysotritia a. ardua. So the other- wise synonymous name of canestrinii should be deleted from the fauna of Switzerland, and all these data henceforth refered to R. a. ardua.

REMARKS ON BORCARD, 1991

Borcard (1991) names five species with species names, but the identified Phthiracarus tardus is accompanied by cf., rendering it uncertain. With high probabil- ity we consider Hoplophthiracarus pavidus (Berlese, 1913) to be H. vanderhammeni Niedbata, 1991. The other three species listed by him (Steganacarus herculeanus, S. striculus and Rhysotritia ardua) are most probably correctly identified.

Concerning his 3 species of Phthiracarus with “open nomenclature” a correct identification is impossible without revision of the material.

458 S. MAHUNKA & L. MAHUNKA-PAPP

REMARKS ON MAHUNKA, 1993a, b

Concerning the 5 published species, only the abandonment of the subgenus Archiphthiracarus has to be mentioned. Its synonymy with Phthiracarus (see Nied- bata, 1992) is here confirmed.

LIST OF THE PUBLISHED SPECIES ACCEPTED FOR THE SWISS FAUNA

Phthiracaridae

Phthiracarus bryobius Jacot, 1936 Phthiracarus crenophilus Willmann, 1951 Phthiracarus globosus (C. L. Koch, 1841)

Steganacaridae

Atropacarus striculus (C. L. Koch, 1835) Atropacarus wandae (Niedbata, 1981) Helvetacarus genavensis Mahunka, 1993a Steganacarus herculeanus Willmann, 1951 Steganacarus magnus (Nicolet, 1855)

Oribotritiidae Mesotritia nuda (Berlese, 1887)

Euphthiracaridae Rhysotritia a. ardua (C. L. Koch, 1841)

LIST OF THE SPECIES TO BE DELETED FROM THE SWISS FAUNA

“Phthiracarus” dasypus (Duges, 1834) “Phthiracarus” piger (Scopoli, 1763) Phthiracarus borealis (Trägärdh, 1910) Phthiracarus laevigatus (C. L. Koch, 1844) Calyptophthiracarus pavidus (Berlese, 1913) Steganacarus clavigerus (Berlese, 1903) Steganacarus spinosus (Sellnick, 1920)

LIST OF LOCALITIES

AG-4 = SWITZERLAND: Argovia: Koblenz, ramification of the river Aar, swamp; 30.IX.1967; leg. C. Besuchet (159).

AG-5 = SWITZERLAND: Argovia: Densbüren, old beech stumps; 4.X.1975; leg. C. Besuchet (160).

AP-1= SWITZERLAND: Appenzell: Hoher Kasten, sifting, 1600-1700m; 18.VIII.1982; leg. C. Besuchet (86).

AP-2 = SWITZERLAND: Appenzell: Santis, 2450m; 10.VII.1967; leg. A. Comellini (161).

AP-4 = SWITZERLAND: Appenzell: Schwägalp, 1400m; 26.VII.2001; leg. C. Besuchet (196).

BE-8 = SWITZERLAND: Berne: Uebischisee near Thun, moss at edge of swamp; 6.VI.1996; leg. C. Besuchet (106).

BE-9 = SWITZERLAND: Berne: Stockhorn, Oberstockensee, mosses at base of rocks, 1700m; 26.V11.1979; leg. C. Besuchet & I. Löbl (165).

BL-1 = SWITZERLAND: Basle-Land: “Reinacher-Heide” near Reinach, Nature Reserve, xerothermic meadows, sifting, 600-700m; 12.X.1989; leg. C. Besuchet (85).

ORIBATIDS FROM SWITZERLAND VII 459

BL-8 = SWITZERLAND: Basle-Land: above Belchen/Eptingen, sifting, 1000m; 29.IX.1979; leg. C. Besuchet (164).

FR-4 = SWITZERLAND: Fribourg: Düdingen, hollow poplar at about 15 metres above ground; 27.1.1983; leg. S. Vit (169).

GE-1 = SWITZERLAND: Geneva: Allondon, Les Baillets, desiccated mosses and lichens; 19.IX.1983; leg. C. Besuchet (58).

GE-2 = SWITZERLAND: Geneva: Bois de Jussy, waterlogged mosses; 30.V1.1983; leg. C. Besuchet (35).

GE-3 = SWITZERLAND: Geneva: Frontenex, bases of oak stumps; 14.VII.1980; leg. C. Besuchet (6).

GE-5 = SWITZERLAND: Geneva: Malval, mosses and grass; 22.X.1982; leg. C. Besuchet (31).

GE-6 = SWITZERLAND: Geneva: Malval, mosses and lichens; 22.X.1982; leg. C. Besuchet (34).

GL-3 = SWITZERLAND: Glarus: Klöntal above Riedern, old tree stumps, 700m; 30.VIIL.1980; leg. C. Besuchet (1).

GR-4 = SWITZERLAND: the Grisons: Felsberg near Chur, dry needles and bark from pine (Pinus sp.) forest; 27.IX.1983; leg. C. Besuchet (57).

GR-5: Switzerland: the Grisons:Swiss National Park, Il Fuorn, under a pile of dead branches, 1800m; 22.IX.1995; leg. C. Besuchet (104).

GR-6 = SWITZERLAND: the Grisons: Landquart, at base of alders (Alnus sp.); 26.IX.1983; leg. C. Besuchet (118).

GR-8 = SWITZERLAND: the Grisons: Samnaun, alpine meadows with Rhododendron sp., sift- ing, 2050m; 26.VIIL.1968; leg. C. Besuchet (26).

GR-9 = SWITZERLAND: the Grisons: Santa Maria Umbrail Pass, sifting, 2000m; 5.VIIL.1974; leg. C. Besuchet (37).

GR-11 = SWITZERLAND: the Grisons: Untervaz near Chur, mosses; 29.IX.1983; leg. C. Besuchet (126).

GR-13 = SWITZERLAND: the Grisons: Val Bregaglia - Val Bondasca, dead leaves and moss- es at base of alders (A/nus sp.), 1400 m; 19.VII—.1984; leg. C. Besuchet (59).

GR-15 = SWITZERLAND: the Grisons: Umbrail Pass, 2000m; 25.VIII.1968; leg. C. Besuchet (149).

GR-16 = SWITZERLAND: the Grisons: Val Bregaglia, Soglio, hollow chestnut stumps, 900m: 10.1X.1985; leg. C. Besuchet (146).

GR-17 = SWITZERLAND: the Grisons: Val Poschiavo, above Cavajone, at base of rocks and Rhododendron sp., 2050-2100m; 27.VII.1983; leg. C. Besuchet (151).

GR-21 = SWITZERLAND: the Grisons: Swiss National Park, Ova Spin, moss at spring-head, 1700m; 28.VII.1929; leg. J. Schweizer (200).

GR-22 = Switzerland: the Grisons: Swiss National Park, S-charl, decaying needles of Mountain Pine (Pinus montana Mill.), 2000m; 24.VII.1032; leg. J. Schweizer (201).

JU-3 = Switzerland: Jura: between Boncourt and Buix, dead leaves; 11.11.1978; leg. C. Besuchet (174).

JU-4 = SWITZERLAND: Jura: Montfaucon, mosses in old peat-bog; 5.VII.2000; leg. C. Besuchet (206).

LU-1 = SWITZERLAND: Lucerne: Eigenthal, peat-bog Forenmoos near the village of Eigenthal, Sphagnum sp., 970m; 2.VIII.1996; leg. C. Besuchet (108).

LU-2 = SWITZERLAND: Lucerne: above Gettnau (between Zell and Willisau), old ant-hill of Formica rufa; 1.VI11.1996; leg. C. Besuchet (107).

LU-5 = SWITZERLAND: Lucerne: Wohlhusen, old tree stumps; 17.11.1979; leg. C. Besuchet (176).

NW-3 = SWITZERLAND: Nidwalden: Trübsee, mosses near spring-head, 1800m; 8.IX.1997; leg. C. Besuchet (127).

460 S. MAHUNKA & L. MAHUNKA-PAPP

SG-8 = SWITZERLAND: St. Gall: Schwendisee above Unterwasser, mosses and grass at lake- side, 1159m; 12.IX.1981; leg. C. Besuchet (179).

SH-1 = SWITZERLAND: Schaffhausen: above Hallau, mosses and dead leaves; 1983; leg. C. Besuchet (50).

SH-3 = SWITZERLAND: Schaffhausen: Bargen, dead leaves: 25.III.1978; leg. C. Besuchet (180).

SO-1 = SWITZERLAND: Solothurn: Ammansegg, mosses on floor of decidous forest; 14.V.1972; leg. S. Mahunka & L. Mahunka-Papp (10).

SO-2 = SWITZERLAND: Solothurn: Ammansegg, litter and dry leaves in mixed forest; 14.V.1972; leg. S. Mahunka & L. Mahunka-Papp (11).

SO-5 = SWITZERLAND: Solothurn: Schnottwil, Bucheggberg, mosses on bark of a live deciduous tree; 27.IX.1987; leg. S. Mahunka & L. Mahunka-Papp (47).

SO-6 = SWITZERLAND: Solothurn: Schnottwil, Bucheggberg, dry needles in pine (Pinus sp.) forest; 27.1X.1987; leg. S. Mahunka & L. Mahunka-Papp (48).

SO-8 = SWITZERLAND: Solothurn: Weissenstein, dead leaves, 1250m; 6.VIII.1979; leg. C. Besuchet (181).

TG-3 = SWITZERLAND: Thurgau: Hudelmoos near Hagenwil, peat-bog with Sphagnum sp., 600m; 13.IX.1993; leg. C. Besuchet (88).

TG-7 = SWITZERLAND: Thurgau: Müllheim, dry leaves in mixed forest; 13.IX.1987; leg. S. Mahunka & L. Mahunka-Papp (42).

TG-8 = SWITZERLAND: Thurgau: Müllheim; rotten trunk with mosses; 13.1X.1987, leg. S. Mahunka & L. Mahunka-Papp (43).

TG-9 = SWITZERLAND: Thurgau: between Bischofszell and Hauptwil, moss on forest floor with ferns; 11.V1.1983; leg. T. & Z. Adamis (21).

TG-10 = SWITZERLAND: Thurgau: between Bischofszell and Hauptwil, moss on forest floor anddry needles in pine (Pinus sp.) forest; 11.V1.1983; leg. T. & Z. Adamis (22).

TG-12 = SWITZERLAND: Thurgau: between Bischofszell and Hauptwil, dry leaves and moss- es on floor in beech forest; 11.V1.1983; leg. T. & Z. Adamis (23).

TI-3 = SWITZERLAND: Ticino: Bordei at base of Mount Gridone (“massif de refuge”), old chestnut stumps in forest, sifting, 700m; 24.IV.1992; leg. C. Besuchet (90).

TI-10 = SWITZERLAND: Ticino: Orselina near Locarno, leaf litter in ravine; 29. VII.1972; leg. C. Besuchet (24).

TI-11 = SWITZERLAND: Ticino: Rancate, chestnut forest, sifting; 7.1X.1965; leg. C. Besuchet (25).

TI-21 = SWITZERLAND: Ticino: Brusino-Arsizio, mosses near rivulet; 20.VII.1990; leg. C. Besuchet (158).

TI-22 = SWITZERLAND: Ticino: Cortascio above Brissago, sifting in ravine, 1050m; 22.1V.1992; leg. C. Besuchet (157).

TI-23 = SWITZERLAND: Ticino: Alpe d’Arena above Vergeletto, at base of rocks, 1700m; 22.V11.1983; leg. C. Besuchet (142).

TI-24 = SWITZERLAND: Ticino: Caprino, compost; 29.V.1982; leg. Besuchet (154).

TI-25 = SWITZERLAND: Ticino: Centovalli, Moneto, dead leaves, 800m; 23.VII.1983; leg. C. Besuchet (144).

TI-26 = SWITZERLAND: Ticino: Valle Onsernone, Spruga, mosses and humus, 1000m; 22.V11.1983; leg. C. Besuchet (141).

TI-28 = SWITZERLAND: Ticino: Rancate, horse-chestnuts; 8.VII.1963; leg. C. Besuchet (183).

TI-32 = SWITZERLAND: Ticino: Valle di Muggio, Bruzella, dead leaves; 3.V1.1969; leg. C. Besuchet (187).

TI-34 = SWITZERLAND: Ticino: Morbio Superiore, tree stump; 6.V.1969; leg. C. Besuchet (Te-69/35) (189).

ORIBATIDS FROM SWITZERLAND VII 461

TI-37 = SWITZERLAND: Ticino: Rancate, old tree stump, extraction by flotation of soil; 5.V1.1969; leg. C. Besuchet & I. Löbl (Te-69/34) (192).

UR-1 = SWITZERLAND: Uri: Klausen Pass, litter of Rhododendron sp., 2000m; 23.VIII.1983;

leg. I. Löbl (60).

UR-2 = SWITZERLAND: Uri: above Urnerboden, dead leaves and dead grass, 1600m;

24.IX.1985; leg. C. Besuchet (147).

UR-3 = SWITZERLAND: Uri: Klausen Pass, sifting under alders, 1950m; 24.VIII.1983; leg.

C. Besuchet (150).

UR-5 = SWITZERLAND: Uri: above Klausen Pass, 2000m; 12.VII.1967; leg. A. Comellini

(194).

UR-6 = SWITZERLAND: Uri: Susten Pass, 2000m; 12.VII.1967; leg. A. Comellini (195).

VD-1b = SWITZERLAND: Vaud: Bonvillars, La Coudre, dead oak branch; 6.VII.1989; leg. J. Steffen (62b).

VD-8 = SWITZERLAND: Vaud: Ollon, old stump; 30.X11.1982; leg. C. Besuchet (56).

VD-9 = SWITZERLAND: Vaud: Vallorbe, source of the river Orbe, roots and soil at base of rocks; 13.V.1982; leg. C. Besuchet & I. Löbl (81).

VS-3 = SWITZERLAND: Valais: Daubensee, mosses and grass, 2200m; 11.VIIL.1980; leg. C. Besuchet (4).

VS-4 = SWITZERLAND: Valais: Daubensee, mosses and grass, 2200m; 11.VII.1980; leg. C. Besuchet (32).

VS-5 = SWITZERLAND: Valais: Fluhalp near Leukerbad, mosses and dead leaves, 2000m; 14. VIII.1980; leg. C. Besuchet (2).

VS-7 = SWITZERLAND: Valais: Finges Forest, pine (Pinus sp.) stumps; 13.VIII.1980; leg. C. Besuchet (36).

VS-12 = SWITZERLAND: Valais: Grand-St-Bernard, mosses on and at base of rocks, 2150m; 10.IX.1996; leg. C. Besuchet (109).

VS-13 = SWITZERLAND: Valais: La Vernaz, waterlogged mosses, 1100m; 4.VI.1985; leg. C. Besuchet (124).

VS-22 = SWITZERLAND: Valais: Simplon Pass, old cow-dung; 2030m; 23.VIII.1997; leg. C. Besuchet (111).

VS-28b = SWITZERLAND: Valais: Vouvry, cave “Grotte de la Pierre à Perret” (VS 13), 490m: 10.VIIL.1989; leg. B. Hauser (63b).

VS-29 = SWITZERLAND: Valais: Randonne above Saillon, under bark, 1100-1350m; 8.X1.1982; leg. C. Besuchet (140).

VS-31 = SWITZERLAND: Valais: Val de Bagnes: Fionnay, mosses and dead leaves: 1450m: 15.V.1990; leg. C. Besuchet (156).

VS-32 = SWITZERLAND: Valais: Zermatt, Riffelberg, mosses, 2500m; 14.VIII.1969; leg. C. Besuchet (148).

VS-33 = SWITZERLAND: Valais: Evolène, mosses in swamp, 1350m; 3.1X.2001; leg. C. Besuchet (198).

VS-34 = SWITZERLAND: Valais: Monthey, outside of the cave “Grotte de Saint-Martin” (VS 1), 540m; 2.V.1980; leg. P. Strinati & V. Aellen (199).

LIST OF NEWLY IDENTIFIED SPECIES (including also the data already published by Mahunka, 1993a, b))

Phthiracaridae Perty, 1841

Helvetacarus genavensis Mahunka, 1993 Locality: GL-3 (Mahunka, 1993a). Distribution: endemic to Switzerland.

462 S. MAHUNKA & L. MAHUNKA-PAPP

Phthiracarus baloghi Feider & Suciu, 1957 Locality: TI-24. Distribution: Near East, SE Europe, Romania, Hungary; first record for Switzerland (the westernmost finding of the species).

Phthiracarus besuchetianus sp. n. Locality: JU-3. Description see below. Phthiracarus boresetosus Jacot, 1930 Localities: AG-5; BL-8; GE-3; GE-5; GR-4; JU-4; SH-3; TG-3. Distribution: Holarctic Region; first record for Switzerland. Phthiracarus bryobius Jacot, 1930 Locality: VS-34 (Mahunka, 1993b). Distribution: HolarcticRegion. Phthiracarus clavatus Parry, 1979 Localities: AP-1; GR-8; JU-4; TI-21; UR-2; VS-22; VS-31. Distribution: Europe; first record for Switzerland. Phthiracarus compressus Jacot, 1930 Localities: AG-4; GL-3; GR-9; JU-3; JU-4; TI-10; VS-4; VS-32. Distribution: Holarctic Region; first record for Switzerland. Phthiracarus crenophilus Willmann,1951 Locality: BL-8. Distribution: Europe. Phthiracarus crinitus (C. L. Koch, 1841) Localities: AG-4; AG-5; LU-5; SH-3; SO-1; TG-3; TG-7; TG-8; TG-9. Distribution: Palaearctic Region, excluding the Far East; first record for Switzerland. Phthiracarus ferrugineus (C. L. Koch, 1841) Localities: AG-4; BE-8; SG-8; SO-8; TI-11; TI-24; TI-25. Distribution: Europe; first record for Switzerland. Phthiracarus globosus (C. L. Koch, 1841) Localities: GE-2; TI-22; UR-2; UR-6; VS-13; VS-28b (Mahunka, 1993b). Distribution: Holarctic Region. Phthiracarus incertus Niedbata, 1983 Locality: SO-8. Distribution: Holarctic Region: first record for Switzerland. Phthiracarus laevigatus (C. L. Koch, 1844) Localities: BL-1; GE-1; GR-4; GR-11; GR-13; GR-15; GR-16; SH-1; SO-1; TI-11; TI- 28; VD-1b. Distribution: Europe; first record for Switzerland. Phthiracarus lentulus (C. L. Koch,1841) Localities: TI-22; TI-28; VS-32. Distribution: Palaearctic Region; first record for Switzerland. Phthiracarus longulus (C.L. Koch, 1841) Localities: FR-4; GR-4; GR-13; LU-5; SH-1; SO-5; SO-6; TG-8; TG-12; TI-10; TI-21; TI-22; TI-25; TI-26; TI-28. Distribution: Holarctic Region: first record for Switzerland. Phthiracarus montanus Pérez-fñigo, 1969 Localities: AP-1; AP-2; SO-8; TI-22; UR-1; VS-3, VS-7. Distribution: Western Palearctic Region; first record for Switzerland. Phthiracarus nitens (Nicolet, 1855) Localities: AG-4; BL-8; JU-3; JU-4; LU-5; SH-3; TG-3. Distribution: Western Palaearctic Region. Phthiracarus opacus Niedbata, 1986 Localities: BL-8; VS-S. Distribution: Palaearctic Region: first record for Switzerland.

ORIBATIDS FROM SWITZERLAND VIII 463

Phthiracarus peristomaticus Willmann, 1951 Locality: GR-13; GR-15; TI-25; TI-26; UR-2; UR-6. Distribution: Palearctic Region; first record for Switzerland. Phthiracarus setosus (Banks, 1885) Locality: TI-32. Distribution: Holarctic Region; first record for Switzerland. Phthiracarus spadix Niedbata, 1983 Localities: AP-1; GR-8; GR-9; GR-13; GR-15; GR-17; NW-3; TI-23; UR-2; UR-3; UR- 5; UR-6; VS-5; VS-22. Distribution: Alps and Carpathian Moutains; first record for Switzerland.

Steganacaridae Niedbata, 1986

Atropacarus csiszarae (Balogh & Mahunka, 1979) Localities: GR-4; TI-22; TI-37. Distribution: Europe, Caucasus; first record for Switzerland. Atropacarus striculus (C. L. Koch, 1835) Localities: BE-8; BE-9; FR-4; GL-3; GR-4; LU-1; LU-5; SG-8; SO-6; TG-3; TI-10; TI- 11; TI-24; TI-25; TI-26; TI-28; TI-37; UR-2; VS-12; VS-22; VS-33. Distribution: Holarctic Region Atropacarus wandae (Niedbata, 1981) Localities: AG-5; JU-3; SH-3; TI-21; VS-34 (Mahunka, 1993b). Distribution: Europe. Calyptophthiracarus c. cretensis (Mahunka, 1979) Locality: TI-22. Distribution: Palearctic Region; first record for Switzerland. Steganacarus (S.) antennatus sp. n. Localities: GR-21; GR-22. Description see below. Steganacarus (S.) applicatus (Sellnick, 1920) Localities: BL-8; GL-3; LU-1; LU-5; TG-7; TG-8; TG-9. Distribution: WesternPalearctic Region; first record for Switzerland. Steganacarus (S.) herculeanus Willmann, 1953 Localities: AG-5; AP-1; AP-4; BL-8; GL-3; LU-5; SO-6; SO-8; VS-4. Distribution: Europe. Steganacarus (S.) hirsutus Pérez-fñigo, 1974 Localities: JU-4; LU-1; SO-1; SO-6; VD-9. Distribution: Western Europe; first record for Switzerland. Steganacarus (S.) magnus (Nicolet, 1855) Localities: JU-3; SH-3. Distribution: Europe. Steganacarus (S.) magnus f. anomalus (Berlese, 1883) Locality: SH-3. Distribution: Palearctic Region; first record for Switzerland. Steganacarus (S.) schweizeri sp. n. Localities: GL-3; LU-5. Description see below. Steganacarus (Tropacarus) carinatus (C. L. Koch, 1841) Localities: GR-11; LU-1; LU-2; TG-3; TI-3; TI-10; TI-22; TI-25. Distribution: Palearctic Region; first record for Switzerland.

Oribotritiidae Grandjean, 1954

Mesotritia nuda Berlese, 1887 Locality: GR-5. Distribution: Europe.

464 S. MAHUNKA & L. MAHUNKA-PAPP

Microtritia minima (Berlese, 1904) Localities: GR-16; LU-2; SH-3; TI-11; TI-22; TI-28; TI-37. Distribution: Europe; first record for Switzerland.

Protoribotritia aberrans Märkel & Meyer, 1959 Locality: GL-3; TI-25. Distribution: Europe; first record for Switzerland.

Protoribotritia oligotricha Märkel, 1963 Locality: GR-5. Distribution: Sweden; first record for Switzerland.

Paratritia baloghi Moritz, 1966 Locality: TI-34. Distribution: Central Europe; first record for Switzerland.

Euphthiracaridae Jacot, 1930

Euphthiracarus cribrarius (Berlese, 1904) Localities: AG-5; GL-3; GR-4; GR-13; JU-4; LU-2; LU-5; SH-3; TG-7; TI-26; VS-29. Distribution: Europe; first record for Switzerland.

Euphthiracarus monodactylus (Willmann, 1919) Localities: AG-4; AG-5; GE-5; GR-6; TI-11; TI-22; TI-37. Distribution: Palearctic Region; first record for Switzerland.

Rhysotritia a. ardua (C. L. Koch, 1841) Localities: AG-4; BL-1; FR-4; GE-1: GE-5: GE-6; GR-4; GR-11; JU-3; SH-1; SH-3; TG-3; TI-11; TI-24; TI-28; TI-37; VD-8: VS-4; VS-29; VS-34 (Mahunka, 1993b). Distribution: Holarctic Region (?).

Rhysotritia ardua hyeroglyphica (Berlese, 1916) Locality: GE-6. Distribution: Europe: first record for Switzerland.

Rhysotritia duplicata (Grandjean, 1953) Localities: LU-2; SO-2; TG-3; TG-9; TG-10; TI-22. Distribution: Europe; first record for Switzerland.

Mesoplophoridae Ewing, 1917

Mesoplophora pulchra Sellnick, 1928 Localities: AG-4; AG-5; GE-3. Distribution: Holarctic Region; first record for Switzerland.

DESCRIPTION AND DISCUSSION OF SOME SPECIES

Phthiracarus baloghi Feider & Suciu, 1957 Figs 1-3

Phthiracarus baloghi Feider & Suciu, 1957: 28, figs 44-55. Phthiracarus baloghi: Niedbata, 1992: 87, pl. 8A-J, 9A-I.

The Swiss specimens differ slightly from the specimens on which the redescription of Niedbata was based. The principal differences occur in the thickness of the sensillus (much longer and narrower on the Swiss specimens, see fig. 3), in the shape of the bothridial squama (significantly longer in the Swiss specimens), and, the proportion of v’ and v” setae on the femur of leg I. The species is also well charac-

ORIBATIDS FROM SWITZERLAND VII 465

Fics 1-3

Phthiracarus baloghi Feider & Suciu, 1957 1: aspis in dorsal view, 2: femur I, 3: aspis in lateral view.

terised by the 1-3 pairs of long c setae, the great length differences of the notogastral setae, as well as the extraordinarily long exobothridial setae. The dimensions of the Swiss specimens fall within the known range.

Phthiracarus besuchetianus sp. n. Figs 4-7

Material examined: Switzerland: Holotype: Jura: JU-3; 13 paratypes: from the same sample. Holotype and 8 paratypes: MHNG? and 5 paratypes (1661-P0-01): HNHM?.

Diagnosis: Sensillus lanceolate, short. 15 pairs of notogastral setae and four pairs of notogastral lyrifissures present. Setae c,, located far from the collar line. The leg chetotaxy is of the “complete type”.

Measurements: Length of aspis: 218-340 um, length of notogaster: 534-608 um, height of notogaster: 388-454 um.

2 MHNG: deposited in the Muséum d’histoire naturelle, Geneva.

3 HNHM: deposited in the Hungarian Natural History Museum, Budapest, with identi- fication number of the specimens in the Collection of Arachnida.

466 S. MAHUNKA & L. MAHUNKA-PAPP

Fics 4-7

Phthiracarus besuchetianus sp. n. 4: body in lateral view, 5: aspis, 6: femur I, 7: anogenital region.

ORIBATIDS FROM SWITZERLAND VIII 467

Aspis: Median crista absent, its outline straight basally, convex anteriorly, with a small concave part rostrally (Fig. 5). Lateral carina distinct, long, reaching over the posterobothridial squama. Lateral rim well developed, sinus line not observable. All prodorsal setae very fine, comparatively short, but setae ex not shorter than the rostral ones. Sensillus lanceolate, with sharply pointed distal end.

Notogaster (Fig. 4): All fifteen pairs of notogastral setae short, fine, filiform, somewhat flagellate at their distal ends. Among setae c,, c, and c, setae c, arise furthest, setae c; nearest to the collar line. Four pairs of lyrifissures present. Vestigial setae of f, located far below setae h,.

Anogenital region (Fig. 7): Typical for the genus, four pairs of long setae arising in the lateral, and five pairs of short setae arising in the medial position. Five pairs of setae present on the anoadanal plates, two of which are much reduced. All short and fine.

Legs: The legs chaetotaxy is of the “complete type”, with setal formula:

1:71 42 241 SAGE 1. IV: 2-1—1—2+1— 10 1:

Setae d of femur I (Fig. 6) located far from the anterior margin of the segment, thick and strongly curved, serrate distal end. Seta /’ on genu IV present.

Remarks: The four pairs of notogastral lyrifissures, the short and lanceolate sensillus and the position of setae c on the notogaster characterize the new species well. It resembles Phthiracarus occultus Niedbata, 1981, but, the new species has 4 pairs of lyrifissures (in occultus only 3), setae c, stand further from the collar line than in occultus, setae d on the femora arise near to the distal margin in occultus (far from the distal margin in the new species) and setae /’ is absent in occultus (present in the new species).

Derivatio nominis: We dedicate the new species to Dr C. Besuchet (Geneva), the renowned coleopterist, for his collecting activity: he gathered the major part of the Swiss soil samples.

Phthiracarus clavatus Parry, 1979

Phthiracarus (Phthiracarus) clavatus Parry, 1979: 338, figs 7a-c, pl. 3: c, f. Phthiracarus (Phthiracarus) clavatus: Niedbata, 1992: 93, pl. 27: a-i, pl. 28: a-e.

It may easily be differentiated from the other species by its long notogastral se- tae, and the characteristically short distance of setae c from the collar line. The species is new for the fauna of Switzerland.

Phthiracarus crinitus (C. L. Koch, 1841) Figs 8-12

Hoplophora crinita C. L. Koch, 1841: f. 32, fig. 8. Phthiracarus crinitus: Kamill, 1981: 263, figs1-7. Phthiracarus crinitus: Niedbata, 1992: 99, pl. 18: a-e, 21: h-l.

The species is readily identified on the basis of the works of Kamill and Niedbata. The Swiss specimens differ somewhat especially from the figure of Kamill, so we also present a figure (see Figs 8-12). The rostral part of the aspis is high, in lateral

468 S. MAHUNKA & L. MAHUNKA-PAPP

Fics 8-10

Phthiracarus crinitus (C. L. Koch, 1841) 8: body in lateral view, 9: bothridial part of the aspis, 10: anogenital region.

view almost appearing cristate. The rostrum projects forward. Seta c, is emitted close to the collar line on the notogaster. Lyrifissure ip sometimes difficult to discern, much weaker than ips. A characteristic feature of the species is that out of the three pairs of setae on the anogenital plate, ad, is not shorter than the anal setae.

The dimensions of the Swiss specimens are within the so far recorded range.

ORIBATIDS FROM SWITZERLAND VIII 469

Fics 11-12 Phthiracarus crinitus (C. L. Koch, 1841) 11: aspis in dorsal view, 12: rostral part of the aspis.

Phthiracarus laevigatus (C. L. Koch, 1841)

Hoplophora laevigata (C. L. Koch, 1841): f. 38, fig. 16. Phthiracarus laevigatus: van der Hammen, 1963: 706, figs 1-6. Phthiracarus laevigatus: Niedbata, 1992: 82, pl 1: a-j.

It is one of the most common species in Switzerland. We identified the species by following the opinion and the redescription of van der Hammen, which had also been accepted by Niedbata (1992). To separate the species of P. laevigatus and P. nitens is not an easy task, further investigation is needed.

Phthiracarus longulus (C. L. Koch, 1841) Figs 13-18

Hoplophora longula C. L. Koch, 1841: f. 32, fig. 17. Phthiracarus longulus: Kamill, 1981: 270, figs 25-31. Phthiracarus longulus: Niedbata, 1992: 116, pl. 71-76.

To identify the species is rather problematic, in spite of the redescriptions prepared by Kamill and Niedbafa, due to the fact that the two redescriptions themselves and the recently collected Swiss specimens somewhat differ from one another. However, the redescriptions and the specimens confirm that the setation of the legs belong to the “reduced” type, the notogaster bears two pairs of lyrifissures, the noto- gastral setae are moderately long, the prodorsal carina is long, the sensillus is lanceo- late and visibly spiculate. The notogaster is slightly elongate and generally light coloured. Kamill defined setae c as arranged in “a row immediately behind the posterior collar margin”, though she depicted them as further away. On the other hand, Niedbata placed them on the collar line, which corresponds well with the situation on the Swiss specimens. However, the latter unequivocally differ from the data of Kamill and Niedbata in the position of the vestigial f,, since both authors place it before /,; the

470 S. MAHUNKA & L. MAHUNKA-PAPP

Fe

x 7] \ 1

Fics 13-15

Phthiracarus longulus (C. L. Koch, 1841) 13: body in lateral view, 14: bothridial part of the aspis, 15: aspis in dorsal view.

Swiss specimens display it behind h, Another difference is the shape of seta d on the femur of leg I; on the Swiss specimens this seta is simple and setiform. For the time being we consider these differences as intraspecific variations only.

Phthiracarus opacus Niedbata, 1986 Figs 19-22

Phthiracarus opacus Niedbata, 1986: Phthiracarus opacus: Niedbata, 1992: 125, pl. 88: A-F.

ORIBATIDS FROM SWITZERLAND VIII 471

Fics 16-18 Phthiracarus longulus (C. L. Koch, 1841) 16: anogenital region, 17: femur I, 18: leg IV.

Our experiences show that the species may be confused with P. peristomaticus. A significant feature is the lack of crista on the aspis. Seta c, on peristomaticus 1s emitted somewhat further from the collar line (Fig. 19) than on opacus. From among the adanal setae ad; is shorter than either the anal or the other two pairs of adanal setae. Niedbata does not mention seta d on the femur of leg I, which is located somewhere in the middle section on opacus, and is apically uncate (Fig. 21), while on peristomaticus the same is a simple, bent seta.

Phthiracarus spadix Niedbata, 1983 Figs 23-28

Phthiracarus spadix Niedbata, 1983: 143, figs 1-17. Phthiracarus spadix: Niedbata, 1992: 136, pl. 101: a-h.

The description of Niedbata corresponds well with the Swiss specimens. Some additional features: on the aspis the lateral keel is often hardly visible. The distal half of sensillus is ornamented with fine spicules. Setae h, and e, clearly bear a few short cilia or spicules. The anterior anal setae are slightly longer, strongly bending forwards and strongly attenuating. From among the genital setae g,-g, are strikingly long, 9-85 somewhat thickened. Seta d is emitted in the middle section of the femur, arcuate, finely ciliate, not reaching the anterior end of this pedal segment.

472 S. MAHUNKA & L. MAHUNKA-PAPP

Fics 19-22

Phthiracarus opacus Niedbata, 1986 19: body in lateral view, 20: aspis in dorsal view, 21: femur I, 22: sensillus.

ORIBATIDS FROM SWITZERLAND VII 473

Fics 23-28

Phthiracarus spadix Niedbata, 1983 23: